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The reduction of information contained in model time series through the use of aggregating statistical performance measures is very high compared to the amount of information that one would like to draw from it for model identification and calibration purposes. It is readily known that this loss imposes important limitations on model identification and -diagnostics and thus constitutes an element of the overall model uncertainty as essentially different model realizations with almost identical performance measures (e.g. r-² or RMSE) can be generated. In three consecutive studies the present work proposes an alternative approach towards hydrological model evaluation based on the application of Self-Organizing Maps (SOM; Kohonen, 2001). The Self-Organizing Map is a type of artificial neural network and unsupervised learning algorithm that is used for clustering, visualization and abstraction of multidimensional data. It maps vectorial input data items with similar patterns onto contiguous locations of a discrete low-dimensional grid of neurons. The iterative training of the SOM causes the neurons to form a discrete, data-compressed representation of the high-dimensional input data. Using appropriate visualization techniques, information on distributions, patterns and relationships in complex data sets can be extracted. Irrespective of their potential, SOM applications have earned very little attention in hydrological modelling compared to other artificial neural network techniques. Therefore, the aim of the present work is to demonstrate that the application of Self-Organizing Maps has very high potential to address fundamental issues of model evaluation: It is shown that the clustering and classification of model time series by means of SOM can provide useful insights into model behaviour. In combination with the diagnostic properties of Signature Indices (Gupta et al., 2008; Yilmaz et al., 2008) SOM provides a novel tool for interpreting the model parameters in the hydrological context and identifying parameter sets that simultaneously meet multiple objectives, even if the corresponding model realizations belong to different models. Moreover, the presented studies and reviews also encourage further studies on the application of SOM in hydrological modelling.
Dry tropical forests undergo massive conversion and degradation processes. This also holds true for the extensive Miombo forests that cover large parts of Southern Africa. While the largest proportional area can be found in Angola, the country still struggles with food shortages, insufficient medical and educational supplies, as well as the ongoing reconstruction of infrastructure after 27 years of civil war. Especially in rural areas, the local population is therefore still heavily dependent on the consumption of natural resources, as well as subsistence agriculture. This leads, on one hand, to large areas of Miombo forests being converted for cultivation purposes, but on the other hand, to degradation processes due to the selective use of forest resources. While forest conversion in south-central rural Angola has already been quantitatively described, information about forest degradation is not yet available. This is due to the history of conflicts and the therewith connected research difficulties, as well as the remote location of this area. We apply an annual time series approach using Landsat data in south-central Angola not only to assess the current degradation status of the Miombo forests, but also to derive past developments reaching back to times of armed conflicts. We use the Disturbance Index based on tasseled cap transformation to exclude external influences like inter-annual variation of rainfall. Based on this time series, linear regression is calculated for forest areas unaffected by conversion, but also for the pre-conversion period of those areas that were used for cultivation purposes during the observation time. Metrics derived from linear regression are used to classify the study area according to their dominant modification processes.rnWe compare our results to MODIS latent integral trends and to further products to derive information on underlying drivers. Around 13% of the Miombo forests are affected by degradation processes, especially along streets, in villages, and close to existing agriculture. However, areas in presumably remote and dense forest areas are also affected to a significant extent. A comparison with MODIS derived fire ignition data shows that they are most likely affected by recurring fires and less by selective timber extraction. We confirm that areas that are used for agriculture are more heavily disturbed by selective use beforehand than those that remain unaffected by conversion. The results can be substantiated by the MODIS latent integral trends and we also show that due to extent and location, the assessment of forest conversion is most likely not sufficient to provide good estimates for the loss of natural resources.
Bei der Mauereidechse (Podarcis muralis, Laurenti 1768) handelt es sich um eine Süd- und mitteleuropäisch verbreitete Art. Sie ist im Süden weitgehend flächendeckend verbreitet, wohingegen sie am Nordrand ihres Verbreitungsgebietes in fragmentierten Populationen vorkommt, deren dauerhaftes Fortbestehen fraglich ist. Das Ziel der vorliegenden Arbeit bestand in der vergleichenden Untersuchung der genetischen Variabilität und Diversität ausgewählter Mauereidechsen-Populationen aus ihrem rezenten mitteleuropäischen Verbreitungsgebiet und ihren Refugialräumen. Weiterhin sollte überprüft werden, ob aus den erhaltenen Daten Rückschlüsse auf eventuell unterschiedliche Einwanderungswege der nördlichen Populationen mittels molekular-genetischer Methoden gezogen werden können. Als Untersuchungsmethode wurde die Mikrosatelliten-DNA Analyse gewählt. Gegenüber der in vorhergehenden Untersuchungen angewandten RAPD-PCR Methode besitzt die Analyse von DNA-Mikrosatelliten den großen Vorteil, über die direkte Bestimmung von Allelfrequenzen wichtige populationsgenetische Daten berechnen zu können. Isoenzymatische Voruntersuchungen an Mauereidechsen erwiesen sich als wenig geeignet. Die SSR-Methode ist besonders gut geeignet, die genetische Diversität und Populationsdifferenzierungen zu ermitteln. Mittels acht spezifischer Mikrosatelliten-Loci wurde die genetische Struktur von sieben Populationen der Mauereidechse untersucht. Als Untersuchungsgebiete wurden Standorte in Italien, Kroatien, Südfrankreich, Nordfrankreich, Rheinland-Pfalz, Nordrhein-Westfalen und Belgien ausgewählt und beprobt. Die untersuchten Loci erwiesen sich alle als polymorph und zum Teil als hoch variabel. Im Rahmen der populationsgenetischen Analysen wurden zahlreiche deskriptive und populationsgenetische Parameter zur Charakterisierung der Populationen verwendet. Wegen unerwarteter Fragmentlängen des Locus A7 wurden exemplarisch einige Allele von etwa 400 bp sequenziert. Dabei konnte zusätzlich außer dem Mikrosatelliten eine etwa 200 bp lange Sequenz nachgewiesen werden, die wahrscheinlich durch eine Insertion entstanden ist. Außerdem wurde ein Basenaustausch (G-C, C-G) beim Vergleich der FOREWARD- und REVERS- Sequenzierung festgestellt. Die Ergebnisse der vorliegenden Arbeit zeigen, dass die Vorkommen der Mauereidechse an ihrem nördlichen Verbreitungsrand teilweise separiert sind. Es konnte gezeigt werden, dass ein Genaustausch der Trierer und der Urfter Tiere untereinander besteht. Eingeschränkt besteht der Genaustausch auch mit den Bitcher Tieren, wohingegen der Genaustausch mit den Belgischen Tieren wohl schon lange eingeschränkt ist oder nur sehr eingeschränkt stattgefunden hat. Die Einwanderung der Mauereidechse in ihr nördliches Verbreitungsgebiet dürfte von den Südfranzösischen Populationen durch das Rhônetal, die Burgundische Pforte, das Rheintal bis zur, in der vorliegenden Arbeit untersuchten Population von Bitche, erfolgt sein. Die Trierer und Urfter Tiere dürften über den Obermoselraum eingewandert sein. Aufgrund der im Verhältnis zu den anderen Populationen stark reduzierten Allelzahl der Population Anhée muss für diesen Standort von einem so genannten bottleneck (Flaschenhals) ausgegangen werden. Der Verdacht, dass sich die zwei Unterarten P. m. brogniardi (Belgien) und P. m. merremia (Nordeifel, Mosel, Bitche) unterscheiden lassen, scheint sich eher zu erhärten, als zu entkräften. Die großen genetischen Unterschiede der Belgischen Population zu den anderen nördlichen Populationen lassen sich am ehesten durch eine Einwanderung über Frankreich an die Maas und einen stark eingeschränkten Genfluss zwischen der Belgischen Population und den restlichen nördlichen Populationen erklären.
This thesis presents a study of tsunami deposits created by the 2004 Indian Ocean tsunami at the Thai Andaman coast. The outcomes of a study are the characteristics of tsunami deposit for paleo-tsunami database, the identification of major sediment layers in tsunami deposit and the reconstructing tsunami run-ups from the characteristics of tsunami deposit for a coastal development program. The investigations of tsunami deposit are made almost 3 years after the event. Field investigations characterize the tsunami deposit as a distinct sediment layer variable in thickness of gray sand deposited with an erosional basis on a pre-existing soil. The best location for the observation of recent tsunami deposit is the area located about 50-200 m inland from the coastline. In most cases, the deposit layer is normally graded. In some cases, the deposit contains rip-up clasts of muddy soils and/or organic matters. The tsunami deposits are compared with three deposits from coastal sub-environments. The mean grain-size and standard deviation of deposits show that the shoreface deposits are fine to very fine sand, poorly to moderately well sorted; the swash zone deposits are coarse to fine sand, poorly to well sorted; the berm/dune deposits are medium to fine sand, poorly to well sorted; and the tsunami deposits are coarse to very fine sand, poorly to moderately well sorted. The plots of deposit mean grain-size versus sorting indicate that the tsunami deposits are composed of shoreface deposits, swash zone deposits and berm/dune deposits as well. The vertical variation of the texture of tsunami deposit shows that the mean grain-size fines upward and fining landward. The analysis and interpretation of the run-up numbers from the characteristics of tsunami deposits get three run-ups for the 2004 Indian Ocean tsunami at the Thai Andaman coast. It corresponds to field observations from the eye-witness reports and local people- affirmations. The total deposition is a major transportation pattern of onshore tsunami sediments. The sediments must fine in the direction of transport. In general, the major origins of the sediment are the swash zone and berm/dune zone where coarse to medium sand is a significant material, the minor origin of tsunami sediment is a shoreface where a significant material is fine to very fine sand. Only at an area with flat slope shorface, the major origin of tsunami sediment is the shoreface. The thicknesses, the mean grain-sizes, and the standard deviations of tsunami deposits are used to evaluate the influences of coastal morphology on the sediment characteristics. The evaluations show that the tsunami affected areas were attacked by the variable energy waves. Wave energies at the direct tsunami wave affected areas are higher than at the indirect tsunami wave affected areas. Tsunami wave energy is highly dissipated at an area with steep slope shoreface. In the same way, tsunami run-up energy is highly dissipated at an area with steep slope onshore. A channel paralleled to the coastline decreases the run-up velocity, slightly dissipates run-up energy. The road and pond highly influence the characteristics of tsunami deposit and tsunami run-up. A road obstructs the run-up velocity, dissipates run-up energy. A pond decreases run-up velocity, dissipates run-up energy. The characteristics of tsunami deposit can be interpreted for reconstructing the characteristics of tsunami run-up such as a run-up height and a flow velocity. Soulsby et al.(2007)- model is applied for reconstructing tsunami run-up at the study areas. The input parameters are sediment grain-size and sediment inundation distance. Ao Kheuy beach and Khuk Khak beach, Phang Nga province, Thailand are the areas listed for reconstructing tsunami run-up. The evaluated run-up heights are 4.2-4.9 m at Ao Kheuy beach, and 5.4-9.4 m at Khuk Khak beach. The evaluated run-up velocities are 12.8-19.2 m/s (maximum) and 0.2-1.9 m/s (mean) at the coastline and onshore, respectively. Hence, a reasonably good agreement between the evaluated and observed run-up is found. Tsunami run-up height and velocity can be used for coastal development and risk management in the tsunami affected areas. The case studies from the Thai Andaman coast suggest that in the area from coastline to about 70-140 m inland was flooded by the high velocity (high energy) run-ups, and those run-up energies were dissipated there. That area ought to be a non-residential area or a physical protection construction area (flood barrier, forest planting, etc.).
This study aims to estimate the cotton yield at the field and regional level via the APSIM/OZCOT crop model, using an optimization-based recalibration approach based on the state variable of the cotton canopy - the leaf area index (LAI), derived from atmospherically corrected Landsat-8 OLI remote sensing images in 2014. First, a local sensitivity and global analysis approach was employed to test the sensitivity of cultivar, soil and agronomic parameters to the dynamics of the LAI. After sensitivity analyses, a series of sensitive parameters were obtained. Then, the APSIM/OZCOT crop model was calibrated by observations over a two-year span (2006-2007) at the Aksu station, combined with these sensitive cultivar parameters and the current understanding of cotton cultivar parameters. Third, the relationship between the observed in-situ LAI and synchronous perpendicular vegetation indices derived from six Landsat-8 OLI images covering the entire growth stage was modelled to generate LAI maps in time and space. Finally, the Particle Swarm Optimization (PSO) and general-purpose optimization approach (based on Nelder-Mead algorithm) were used to recalibrate four sensitive agronomic parameters (row spacing, sowing density per row, irrigation amount and total fertilization) according to the minimization of the root-mean-square deviation (RMSE) between the simulated LAI from the APSIM/OZCOT model and retrieved LAI from Landsat-8 OLI remote sensing images. After the recalibration, the best simulated results compared with observed cotton yield were obtained. The results showed that: (1) FRUDD, FLAI and DDISQ were the major cultivar parameters suitable for calibrating the cotton cultivar. (2) After the calibration, the simulated LAI performed well with an RMSE and mean absolute error (MAE) of 0.45 and 0.33, respectively, in 2006 and 0.46 and 0.41, respectively, in 2007. The coefficient of determination between the observed and simulated LAI was 0.83 and 0.97, respectively, in 2006 and 2007. The Pearson- correlation coefficient was 0.913 and 0.988 in 2006 and 2007, respectively, with a significant positive correlation between the simulated and observed LAI. The difference between the observed and simulated yield was 776.72 kg/ha and 259.98 kg/ha in 2006 and 2007, respectively. (3) Cotton cultivation in 2014 was obtained using three Landsat-8 OLI images - DOY136 (May), DOY 168 (June) and DOY 200 (July) - based on the phenological differences in cotton and other vegetation types. (4) The yield estimation after the assimilation closely approximated the field-observed values, and the coefficient of determination was as high as 0.82, after recalibration of the APSIM/OZCOT model for ten cotton fields. The difference between the observed and assimilated yields for the ten fields ranged from 18.2 to 939.7 kg/ha. The RMSE and MAE between the assimilated and observed yield was 417.5 and 303.1 kg/ha, respectively. These findings provide scientific evidence for the feasibility of coupled remote sensing and APSIM/OZCOT model at the field level. (5) Upscaling from field level to regional level, the assimilation algorithm and scheme are both especially important. Although the PSO method is very efficient, the computational efficiency is also the shortcoming of the assimilation strategy on a regional scale. Comparisons between the PSO and general-purpose optimization method (based on the Nelder-Mead algorithm) were implemented from the RSME, LAI curve and computational time. The general-purpose optimization method (based on the Nelder-Mead algorithm) was used for the regional assimilation between remote sensing and the APSIM/OZCOT model. Meanwhile, the basic unit for regional assimilation was also determined as cotton field rather than pixel. Moreover, the crop growth simulation was also divided into two phases (vegetative growth and reproductive growth) for regional assimilation. (6) The regional assimilation at the vegetative growth stage between the remote sensing derived and APSIM/OZCOT model-simulated LAI was implemented by adjusting two parameters: row spacing and sowing density per row. The results showed that the sowing density of cotton was higher in the southern part than in the northern part of the study area. The spatial pattern of cotton density was also consistent with the reclamation from 2001 to 2013. Cotton fields after early reclamation were mainly located in the southern part while the recent reclamation was located in the northern part. Poor soil quality, lack of irrigation facilities and woodland belts of cotton fields in the northern part caused the low density of cotton. Regarding the row spacing, the northern part was larger than the southern part due to the variation of two agronomic modes from military and private companies. (7) The irrigation and fertilization amount were both used as key parameters to be adjusted for regional assimilation during the reproductive growth period. The result showed that the irrigation per time ranged from 58.14 to 89.99 mm in the study area. The spatial distribution of the irrigation amount is higher in the northern part while lower in southern study area. The application of urea fertilization ranged from 500.35 to 1598.59 kg/ha in the study area. The spatial distribution of fertilization was lower in the northern part and higher in the southern part. More fertilization applied in the southern study area aims to increase the boll weight and number for pursuing higher yields of cotton. The frequency of the RSME during the second assimilation was mainly located in the range of 0.4-0.6 m2/m2. The estimated cotton yield ranged from 1489 to 8895 kg/ha. The spatial distribution of the estimated yield is also higher in the southern part than the northern study area.
Die Zweckbestimmung der modernen Clustertheorie liegt in der Analyse regional-sektoraler Verbundsysteme von ökonomischen Aktivitäten. Erst seit kurzem wird der Versuch einer Übertragung auf touristische Destinationen in Entwicklungsländern unternommen. Die vorliegende Dissertation untersucht theoretische, methodische und praktische Implikationen dieser Übertragung anhand des Fallbeispiels der touristischen Clusterentwicklung in der Napo-Provinz Ecuadors. Hierbei stehen insbesondere die Rahmenbedingungen für die Genese wissensbasierter (kompetitiver) Wettbewerbsvorteile als Quelle der Clusterdynamik im Zentrum. Anhand einer quantitativen Methodik wird in diesem Rahmen sowohl ein Nachweis zur Gültigkeit eines vorab abgeleiteten Clusterentwicklungsmodells geführt, als auch auf dieser Grundlage ein Beitrag zur Verbesserung bestehender Diagnosesysteme geleistet.
The state-of-the-art finite element software Plaxis 3D was applied in a real-world study site of the Turaida castle mound to investigate the slope stability of the mound and understand the mechanisms triggering landslides there. During the simulation, the stability of the castle mound was analysed and the most landslide-susceptible zones of hillslopes were determined. The 3D finite-element stability analysis has significant advantages over conventional 2D limit-equilibrium methods where locations of 2D stability sections are arbitrarily selected. Two modelling scenarios of the slope stability were elaborated considering deep-seated slides in bedrock and shallow landslides in the colluvial material of slopes. The model shows that shallow slides in colluvium are more probable. In the finite-element model, slope failure occurs along the weakest zone in colluvium, similarly to the situation observed in previous landslides in the study site. The physical basis of the model allows results to be obtained very close to natural conditions and delivers valuable insight in triggering mechanisms of landslides.
Laboratory landslide experiments enable the observation of specific properties of these natural hazards. However, these observations are limited by traditional techniques: frequently used high-speed video analysis and wired sensors (e.g. displacement). These techniques lead to the drawback that either only the surface and 2D profiles can be observed or wires confine the motion behaviour. In contrast, an unconfined observation of the total spatiotemporal dynamics of landslides is needed for an adequate understanding of these natural hazards.
The present study introduces an autonomous and wireless probe to characterize motion features of single clasts within laboratory-scale landslides. The Smartstone probe is based on an inertial measurement unit (IMU) and records acceleration and rotation at a sampling rate of 100 Hz. The recording ranges are ±16 g (accelerometer) and ±2000∘ s−1 (gyroscope). The plastic tube housing is 55 mm long with a diameter of 10 mm. The probe is controlled, and data are read out via active radio frequency identification (active RFID) technology. Due to this technique, the probe works under low-power conditions, enabling the use of small button cell batteries and minimizing its size.
Using the Smartstone probe, the motion of single clasts (gravel size, median particle diameter d50 of 42 mm) within approx. 520 kg of a uniformly graded pebble material was observed in a laboratory experiment. Single pebbles were equipped with probes and placed embedded and superficially in or on the material. In a first analysis step, the data of one pebble are interpreted qualitatively, allowing for the determination of different transport modes, such as translation, rotation and saltation. In a second step, the motion is quantified by means of derived movement characteristics: the analysed pebble moves mainly in the vertical direction during the first motion phase with a maximal vertical velocity of approx. 1.7 m s−1. A strong acceleration peak of approx. 36 m s−2 is interpreted as a pronounced hit and leads to a complex rotational-motion pattern. In a third step, displacement is derived and amounts to approx. 1.0 m in the vertical direction. The deviation compared to laser distance measurements was approx. −10 %. Furthermore, a full 3D spatiotemporal trajectory of the pebble is reconstructed and visualized supporting the interpretations. Finally, it is demonstrated that multiple pebbles can be analysed simultaneously within one experiment. Compared to other observation methods Smartstone probes allow for the quantification of internal movement characteristics and, consequently, a motion sampling in landslide experiments.
The fragmentation of landscapes has an important impact on the conservation of biodiversity. The genetic diversity is an important factor for a population- viability, influenced by the landscape structure. However, different species with differing ecological demands react rather differently on the same landscape pattern. To address this feature, we studied ten xerothermophilous butterfly species with differing habitat requirements (habitat specialists with low dispersal power in contrast to habitat generalists with low dispersal power and habitat generalists with higher dispersal power). We analysed allozyme loci for about 10 populations (Ã 40 individuals) of each species in a western German study region with adjoining areas in Luxemburg and north-eastern France. The genetic diversity and genetic differentiation between local populations was discussed under conservation genetic aspects. For generalists we detected a more or less panmictic structure and for species with lower abundance and sedentarily behaviour the effect of isolation by distance. On the other hand, the isolation of specialists was mostly reflected by strong genetic differentiation patterns between the investigated populations. Parameters of genetic diversity were mostly significantly higher in generalists, compared to specialists. Substructures within populations as an answer of low intrapatch migration, low population densities and high population fluctuations could be shown as well. Aspects of landscape history (the historical distribution of habitats resulting of the presence of limestone areas) and the changes of extensive sheep pasturing and the loss of potential habitats in the last few decades (recent fragmentation) are discussed against the gained genetic data-set of the ten butterflies.
As an interface between an individual and its environment, the skin is a major site of direct exposure to exogenous substances. Once absorbed, these substances may interact with different biomolecules within the skin. The aryl hydrocarbon receptor (AhR) signaling pathway is one mechanism whereby the skin responds to exposures, predominantly through the induction or upregulation of metabolizing enzymes. One known physiological role of the AhR in many tissues is its involvement in the control of cell cycle progression. In skin, almost nothing is known about this physiological function. Moreover, the question whether frequently used naturally occurring phenolic derivatives like eugenol and isoeugenol impact on the AhR within the skin has rarely been studied so far. Eugenol and isoeugenol are due to their odour referred to as fragrances. The ubiquitous distribution of eugenol and isoeugenol results in an almost unavoidable contact with these substances in our daily lives. Despite this fact, their molecular mechanisms of action in skin are poorly understood. There is evidence supporting the hypothesis that these substances may impact on the AhR. On the one hand, eugenol is shown to induce cytochrome P450 1A1 (CYP1A1), a well-known target gene of the AhR. On the other hand, their known anti-proliferative properties might also be mediated by the AhR, based on its physiological function. In order to proof this hypothesis, it was investigated whether eugenol and isoeugenol impact on the AhR signaling pathway in skin cells. Results revealed that eugenol as well as isoeugenol impact on the AhR signaling pathway in skin cells. Both substances caused the translocation of the AhR into the nucleus, induced the expression of the well-known AhR target genes CYP1A1 and AhR repressor (AhRR) and exhibited impact on cell cycle progression. Both substances caused an AhR-dependent cell cycle arrest in skin cells, modulated protein levels of several cell cycle regulatory proteins, inhibited DNA synthesis and thereby reduced cell numbers. The comparison of wildtype cells to AhR knockdown cells revealed an influence of the AhR on cell cycle progression in skin cells in the absence of exogenous ligands. AhR knockdown cells exhibited a slower progression through the cell cycle caused by an accumulation of cells in the G0/G1 phase of the cell cycle and a decreased DNA synthesis rate. Modulation of cell cycle regulatory proteins involved in the transition from the G0/G1 to the S phase of the cell cycle was altered in AhR knockdown cells as well. To conclude, eugenol as well as isoeugenol were able to impact on the AhR signaling pathway in skin cells. Their molecular mechanisms of action are similar to those of classical AhR ligands, although their structural characteristics strongly differ from that of these ligands. In the absence of exogenous ligands the AhR promotes cell cycle progression in many tissues and this knowledge could be expanded on skin-derived cells within the scope of this thesis.